The story revolves around Charles Karino, a Levamme Empire mercenary aerial pilot who mans the twin-seater reconnaissance seaplane Santa Cruz. One day, he receives a preposterous assignment: to fly solo over 12,000 kilometers of enemy waters to transport a girl named Fana del Moral. Fana happens to be betrothed empire's crown prince and is a girl possessing beauty "equal to 5,000 beams of light".
 
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